Upon arrival in San José, the capital of Costa Rica, transfer by land to Puerto Caldera and embark on the National Geographic Quest. Enjoy a sunset dinner onboard as you chart a course for the Guanacaste region on Costa Rica’s Pacific coast.

Drop anchor off the remote Papagayo Peninsula, where you’ll spend the day soaking up the sun and enjoying beach time alongside waters fringed by offshore islets, which create dynamic habitat for a variety of tropical fish. Or choose to walk along a rarely used four-wheel drive track, which takes you back into a beautiful gallery forest. The area is rich with bird and plant varieties, and the naturalists describe the intricate web of life in these oceanside ecosystems.

After lunch on board, you continue to explore the coastline of Bahía Huevos and Playa Celestia. The Zodiacs take you on an exploration of the bay and the peninsula. If conditions permit, you drop kayaks and paddleboards and explore the nearby mangroves.

Weather permitting, don a snorkel and fins to discover a rich underwater world or hike into the upper reaches of the Guanacaste Conservation Area for commanding views over the full length of this rarely visited archipelago.

Back on board, you sail toward the mainland while enjoying a delicious lunch. Depending on the conditions, you may choose to explore Santa Elena Bay in Santa Rosa National Park, Junquillal Wildlife Refuge, or Cuajiniquil Bay, all part of the extraordinarily biodiverse Guanacaste Conservation Area. While venturing out by Zodiac, you immerse yourself in the tropical environment, alive with color. Yellow and black great kiskadee sally from exposed perches looking to capture a flying insect for an evening meal, and orange-fronted parakeets chatter overhead. Predators such as mangrove hawks may take wind, setting out on a late afternoon hunt, while ringed kingfishers, richly colored in brick red and deep blue, wait patiently, watching the shallows for a fish’s silvery flash.

The adventure today continues as you set off inland to explore Rincón de la Vieja National Park. This mountainous region is also a part of the Guanacaste Conservation Area and encompasses more than 34,000 acres. The area, with its forests carpeting the sides of two active volcanoes, is alive with geothermal activity as gurgling, bubbling mud pots spout and steam rises above the canopy. Choose between several enticing options today including a hike through the transition zone where tropical dry habitat meets the temperate cloud forest. Here, you keep an eye out for exotic wildlife, with the possibility of encountering mantled howler monkeys and regionally special birds including the noisy and conspicuous white-throated magpie-jay. Alternatively, after a short hike into the forest, opt to swim below Las Chorreras Waterfall or soar through the canopy by zipline. Afterwards, enjoy a sumptuous lunch served poolside at Hacienda Guachipelin, including fresh ceviche.

Along the coast of the Nicoya Peninsula, and in many ways demarcating the line between the wetter jungles of the south and Guanacaste’s drier environment, sits Curú National Wildlife Refuge, Costa Rica’s first private national wildlife reserve. Learn about the hundreds of species of birds, reptiles, and mammals that inhabit this thriving ecosystem while exploring the park’s tracks and trails. Here you look for white-faced capuchin monkeys, turquoise-browed motmots, and with luck, scarlet macaws. The northern tamandua resides here, and while often elusive and quiet, this primarily arboreal member of the anteater family may be spotted.

Later, as you enjoy lunch on board, you cruise to a cluster of offshore islets collectively called Isla Tortuga. Relax on the beach shaded by swaying palms and tropical vegetation, then celebrate the finale of the Guanacaste adventure with afternoon refreshments and hors d'oeuvres on the beach.

Disembark after breakfast in Puerto Caldera. Drive to the airport in San José for flights home.